Aria Montgomery is portrayed sa pamamagitan ng Lucy Hale. She moves back to Rosewood with her mother, Ella, father, Byron, and younger brother, Mike, after a taon long absence in Iceland. She is secretly tormented after discovering her father having an affair with one of his students. After meeting a man named Ezra Fitz and discovering that he's her English teacher, Aria starts a secret affair with him. Aria and Ezra struggle with the forbidden nature of their relationship as Aria is tormented sa pamamagitan ng keeping the secret of her father's past affair from her mother. Just as Aria decides to tell the truth, she finds that "A" has beaten her to it in a tell-all letter to Ella.
Hanna Marin is portrayed sa pamamagitan ng Ashley Benson. She was Alison's lackey and has taken Alison's place as the "It Girl," along with former loser, Mona Vanderwaal. Alison used to constantly tease Hanna about her weight, which in turn made Hanna bulimic, and is something "A" taunts Hanna about. Hanna shoplifts for attention from her workaholic mother, Ashley, and her father, who left her and her mother a taon ago, along with Mona. When Hanna is caught, Hanna's image-conscious mother sleeps with the police officer, Darren Wilden, to get the charges drop and avoid scandal. Hanna's chastity boyfriend, Sean, vows his virginity, much to Hanna's anger, which she takes out sa pamamagitan ng stealing and crashing Sean's car. And suddenly, Hanna's absent father returns with a fiancée and a seemingly perfect stepdaughter and forces Hanna to take responsibility sa pamamagitan ng getting a job at Sean's mother's office.
Spencer Hastings is portrayed sa pamamagitan ng Troian Bellisario. Spencer is smart, pretty, and always is in the shadows of her smarter and prettier older sister, Melissa, who's newly-engaged to a British hottie, Wren, who seems to have madami of an interest in Spencer than in Melissa. Spencer finds herself increasingly attracted to her sister's fiancé, Wren, whom "A" warns Spencer not to halik or "A" will tell. Spencer and Wren do halik and are caught sa pamamagitan ng Melissa, who insists that Wren moves out. Spencer has trouble Pagsulat a paper for class because Melissa and their parents are mad at her, so Spencer submits one of Melissa's old papers as her own. Spencer is horrified when her paper is nominated for a national award.
Emily Fields is portrayed sa pamamagitan ng Shay Mitchell. Emily is a competitive swimmer who comes from a conservative family. She befriends Maya St. Germain, the new resident of Alison's former house. Maya introduces Emily to marijuana, and the girls grow closer. A message from "A" reveals Emily to be curious about her sexuality, and hints that she had kissed Alison in the past. Emily's mother invites Maya to stay for a while, and she and Emily share a room, in which they also share a bed. Emily's boyfriend becomes possessive and almost forces her to have sex with him, but she is saved sa pamamagitan ng Toby. Emily breaks up with her boyfriend and kisses Maya in in a litrato booth at a party; someone steals the photograph, which she later finds in her textbook. Emily starts to befriend Toby, a former target sa pamamagitan ng Alison.
Alison DiLaurentis is played sa pamamagitan ng Sasha Pieterse. Alison is the former "It Girl" of Rosewood and the leader of the clique. Ruling the group with an iron fist, the somewhat vicious Alison had gone missing during a slumber party; a taon later, her disappearance is still unsolved. When Aria, Hanna, Spencer, and Emily begin receiving threatening messages from "A" reminding them of their darkest secrets, they suspect that "A" is Alison, whose body is soon discovered buried in her former backyard.
Ezra Fitz, or Mr. Fitz, is portrayed sa pamamagitan ng Ian Harding, the new English teacher in Rosewood who becomes romantically involved with Aria the araw before she appears in his class. They try to resist their attraction, but fail.
Ella and Byron Montgomery
Ella (Holly Marie Combs) and Byron (Chad Lowe) are the parents of Aria and Mike Montgomery. Ella is the kind, caring stay-at-home mother and Byron is the adulterous history professor who cheated on his wife a taon nakaraan with his student, Meredith. Ella unknowingly befriends Meredith. Byron regrets the affair and Aria struggles with keeping the secret from her mother to protect her. The relationship between Byron and Ella is strained after Ella reads a letter from "A" revealing Byron's affair.
Ashley Marin, played sa pamamagitan ng Laura Leighton, Hanna's workaholic mother who is raising her daughter on her own after her husband left her. Abandoned sa pamamagitan ng Hanna's father, Ashley recognizes Hanna's wild behavior as a cry for attention. Warning her daughter that any indiscretion in their small town can humiliate them, Ashley avoids Hanna being charged for shoplifting sa pamamagitan ng sleeping with the arresting officer.
Melissa is Spencer's older sister who is engaged to Wren. She is prettier and smarter than Spencer and tends to be her parents favorite. She sees Spencer halik Wren and insists that he moves out. She is played sa pamamagitan ng Torrey DeVitto.
Maya St. Germain
Bianca Lawson plays Maya, whose family has moved into Alison's old house. She and Emily bond over some of Alison's discarded belongings and Maya's marijuana, and Maya's presence awakens a slight same-sex attraction in Emily. The discovery of Alison's body buried under the gazebo on Maya's property brings the girls closer. They halik at the party, but Emily starts to push Maya away when Emily reengages with Toby Cavanaugh.
Janel Parrish portrays Mona Vanderwaal, a former social outcast who longed to be accepted into Alison's group. In the taon since Alison's disappearance, Mona has transformed herself into a popular social leader alongside Hanna.
Known as Jenna Cavanaugh in the book series, Jenna is portrayed sa pamamagitan ng Tammin Sursok, the younger stepsister of Toby, Jenna is the girl who Nawawala her sight due to a prank played sa pamamagitan ng Alison, intended for Toby. Jenna is in psychotherapy. The girls suspect her to be the mysterious "A".
Keegan Allen portrays Toby Cavanaugh, Jenna's stepbrother. Ali's prank was meant for him, but ended up hurting Jenna; he knows the girls were behind it, but took the blame when Ali threatened to reveal one of his secrets. He is attracted to Emily and they start hanging out.
Sean is Hanna's boyfriend who pledges his virginity. Hanna's anger and embarrassment at Sean's refusal to have sex makes Hanna take Sean's car for a joyride and ultimately crash it. Sean forgives Hanna and reveals that he is a member of RLW, Real pag-ibig Waits, which is a virginity club. He is played sa pamamagitan ng Chuck Hittinger.
Wren is portrayed sa pamamagitan ng Julian Morris. Wren was engaged to Melissa until he develops feelings for Spencer. When he was caught halik with Spencer sa pamamagitan ng Melissa, she calls off the engagement and kicks Wren out of their barn. He then apologizes to Spencer for making a mess with her and her sister and told her that he wishes that he met her first.